Senior lecturer


  • Teaching and research activities
  • Supervision of PhD and Master students
  • Tutor of the virtual campus Do It Your Soil

A blog on earthworms:

Teaching activities
Given lessons

Useful links for teachers

Research topics

  • Role of soil organic matter and soil fauna in pedogenetic processes, regarding especially soil structure
  • Soil biological activities (earthworms, microorganisms) and their influence on the redistribution, availability and transfer of soil nutrients. Relationships between root systems and plant growth
  • Ecosystems studied: agricultural plots, permanent pastures and alluvial ecosystems. Use of experimental designs as microcosms and rhizoboxes


Amossé J., Jelmini J.-P., Havlicek E., Mitchel E.A.D., Le Bayon R.-C., Gobat J.-M. (2014). Mille ans d’extension urbaine à Neuchâtel : évolution des paysages et des sols. Bulletin de la SNSN, 134: 45-66.

Amossé, J., Le Bayon R.-C., Gobat, J.-M., (2014). Are urban soils similar to natural soils of river valleys? Journal of Soils and Sediments. Special Issue: SUITMA 7: DOI 10.1007/s11368-014-0973-6.

Le Bayon R.-C., Bullinger-Weber G., Gobat J.-M., Guenat C. (2013). Earthworm communities as indicators for evaluating floodplain restoration success. In Environmental Management, Restoration and Ecological Implications, 47-68 pp.

Gobat J.-M., Le Bayon R.C. (2013). Structure des sols et êtres vivants. In Les sols et leurs structures - Observations à différentes échelles. Eds: Baize D., Duval O., Richard G. QUAE Editions, INRA, Paris.

Kohler-Milleret R., Le Bayon R.C., Chenu C., Gobat J.-M., Boivin P. (2013). Impact of two root systems, earthworms and mycorrhizae ont the physical properties of an unstable silt loam Luvisol and plant production. Plant and Soil, 370: 251-265.

Jabiol B., Zanella A., Ponge J.-F., Sartori G., Englisch M., Van Delft B., De Waal R., Le Bayon R.C. (2013). A proposal for including humus forms in the World Reference Base for Soil Resources (WRB-FAO). Geoderma, 192:  286-294.

Fournier B., Samaritani E, Shrestha J., Mitchell E.A.D. Le Bayon R.C. (2012). Community ecology of earthworms in a restored floodplain and potential as bioindicators of river restoration. Applied Soil Ecology, 59: 87-95.

Bullinger-Weber G., Guenat C., Salomé C., Gobat J.-M., Le Bayon R.C. (2012). Impact of flood deposits on earthworm communities in alder forests from a subalpine floodplain (Kandersteg, Switzerland). European Journal of Soil Biology, 49: 5-11.

Le Bayon R.C., Matera V., Kohler-Milleret R., Degen C., Gobat J.-M. (2011). Earthworm activity alters geogenic arsenic and soil nutrient dynamics. Pedobiologia, 54S: S193-S201.

Salomé C., Guenat C., Bullinger-Weber G., Gobat J.-M., Le Bayon R.C. (2011). Earthworm communities in alluvial forests: influence of altitude, vegetation stages and soil parameters. Pedobiologia, 54S: S89-S98.

Chapuis-Lardy L., Le Bayon R.C., Brossard M., Lopez-Hernandez D., Blanchart E. (2011). Role of soil macrofauna in P cycling. In "Phosphorus in Action - Biological Processes in Soil Phosphorus Cycling", Bünemann E.K., Oberson A. and Frossard E. (Eds). Springer Soil Biology Series 26, Springer, NY, USA, pp. 199-213.

Milleret R., Le Bayon R.C., Tarnawski S., Boivin P., Gobat J.-M. (2010). Earthworm, mycorrhiza and root interactions: their effects on some chemical, physical and biological soil properties. Bulletin de la Société Suisse de Pédologie, 30: 81-83.

Le Bayon R.C., Milleret R. (2009) Effects of earthworms on phosphorus dynamics - a review. Dynamic Plant, Dynamic Soil 3 (Special issue 2): 21-27.

Milleret R., Le Bayon R.C., Lamy F., Gobat J.-M., Boivin P. (2009) Impact of root, mycorrhiza and earthworm on soil physical properties as assessed by shrinkage analysis. Journal of Hydrology, 373: 499-507.

Weisskopf L., Akello P., Milleret M., Khan Z.R., Schulthess F., Gobat J.-M., Le Bayon R.C. (2009) White lupin increases the yield of maize when used as a companion crop, as a cover crop and as a source of organic matter. Plant and Soil, 319:101-114.

Milleret R., Le Bayon R.C., Gobat J.-M. (2009) Root, mycorrhiza and earthworm interactions: their effects on soil structuring processes, plant and soil nutrient concentration and plant biomass. Plant and Soil, 316:1-12.

Weisskopf L.,Le Bayon R.C., Kohler F., Page V., Jossi M., Gobat J.-M., Martinoia E., Aragno M. (2008) The impact of plant species and organic acid exudation on rhizosphere bacterial communities, as revealed by a long term comparative study on lupin and wheat. Soil Biology and Biochemistry, 40: 1772-1780.

Degen C.,Le Bayon R.C. (2008) Activité biologique dans le sol agricole : ça grouille sous nos pieds! Rubrique "Rendez-vous nature". Agri, Hebdomadaire professionnel agricole de la Suisse romande.

Bütler R., Lita Patty L., Le Bayon R.C., Guenat C., Schlaepfer R. (2007) Log decay of Picea abies in the Swiss Jura Mountains of central Europe. Forest Ecology and Management, 242: 791-799.

Bullinger-Weber G.,Le Bayon R.C., Guenat C, Gobat J.-M. (2007) Influence of some physicochemical and biological parameters on soil structure formation in alluvial soils. European Journal of Soil Biology, 43: 57-70.

Page V., Le Bayon R.C., Feller U. (2006) Partitioning of zinc, cadmium, manganese and cobalt in wheat (Triticum aestivum) and lupin (Lupinus albus) and further release into the soil. Environmental and Experimental Botany, 58: 269-278

Binet F., Kersanté A., Munier-Lamy C., Le Bayon R.C., Belgy M.J, Shipitalo M.J. (2006) Lumbricid macrofauna alter atrazine mineralization and sorption in a silt loam soil. Soil Biology and Biochemistry, 38: 1255-1263.

Le Bayon R.C., Jansa J., Frossard E. (2006b) A method sheet on microcosms. In: COST 631 Handbook of methods used in rhizosphere research. Chapter 1.1: Growth systems. Lüster, J. And Finlay, R. (Eds). Pp. 141-142.

Le Bayon R.C., Weisskopf L., Martinoia E., Jansa J., Frossard E., Keller F., Föllmi K.B. and Gobat J.-M. (2006a) Soil phosphorus uptake by continuously cropped Lupinus albus. A new microcosm design. Plant and Soil, 283: 309-321.

Le Bayon R.C., Binet F. (2006) Earthworms change the distribution and availability of phosphorous in organic substrates. Soil Biology and Biochemistry, 38: 235-246.

Shipitalo M.J., Le Bayon R.C. (2004) - Quantifying the effects of earthworms on soil aggregation and porosity. In Edwards C.A. (Ed.), Earthworm Ecology, 2nd Edition , CRC Press LLC, pp. 183-200.

Le Bayon R.C., Moreau S., Gascuel-Odoux C., Binet F. (2002) - Annual variations in earthworm surface-casting activity and soil transport by water runoff under a temperate maize agroecosystem. Geoderma, 106: 121-135.

Le Bayon R.C., Binet F. (2001) - Earthworm surface casts affect soil erosion by runoff water and phosphorus transfer in a temperate maize crop. Pedobiologia, 45: 430-442.

Binet F., Le Bayon R.C. (1999) - Space-time dynamics in situ of earthworm casts under temperate cultivated soils. Soil Biology and Biochemistry, 31: 85-93. F

Le Bayon R.C., Binet F. (1999) - Rainfall effects on erosion of earthworm casts and phosphorus transfer by water runoff. Biology and Fertility of Soils, 30: 7-13.

Le Bayon R.C. (1999) -  Influence des activités des lombriciens sur la dynamique (disponibilité, transfert) du phosphore en sols tempérés. Thèse de doctorat, Université de Rennes 1 (France).